Gillian pays a heavy price for her disloyalty to Diana, but not before news that vampires are studying witch DNA reaches Knox. The fall-out threatens the fragile equilibrium between creatures and the focus is on ending Matthew and Diana’s relationship whatever the cost. Agatha treads a careful line of democracy at the Congregation, but Nathaniel continues to instigate daemon direct-action. Ysabeau relies on shock tactics to expel Diana until Matthew returns with breathtaking news about Diana’s DNA. Diana and Matthew are finally united in love, only for the unexpected to happen